- Scientific name
- Pachycudonia spathulata
- Author
- (S. Imai) S. Imai
- Common names
-
- IUCN Specialist Group
Cup-fungi, Truffles and Allies
- Kingdom
- Fungi
- Phylum
- Ascomycota
- Class
- Leotiomycetes
- Order
- Helotiales
- Family
- Incertae sedis
- Assessment status
-
Published
- Assessment date
- 2024-12-15
- IUCN Red List Category
-
LC
- Assessors
- Noah Siegel (25 Prospect Hill Road, Royalston, MA 01368, US / Global Fungus Assessment); Gregory Mueller (Chicago Botanic Garden / IUCN SSC Mushroom, Bracket and Puffball Specialist Group)
- Reviewers
- David Minter (CABI International, UK / IUCN SSC Cup-fungus, Truffle and Ally Specialist Group)
Assessment Notes
Justification
Pachycudonia spathulata is a small fungus with an irregularly rounded, convoluted to somewhat spatulate, bright golden yellow to orange ‘head’, a pale orange stipe that becomes brownish from the base upwards in age, and leathery, non-gelatinous flesh. Most reports are from winter and early spring from under
Arctostaphylos spp.
It remains a poorly known species, but was recently included in "A Field Guide to the Rare Fungi of California’s National Forests" (Siegel
et al. 2019) and "FunDiS Rare 10 Challenge" (Fungal Diversity 2020).
Based on habitat and current reports, this species is likely to be far more common than records indicate. It is assessed as Least Concern.
Taxonomic notes
Described as
Cudonia spathulata based on a 1931 collection made in Big Basin State Park in Santa Cruz County, California, USA (Imai 1942), and later transferred into the genus
Pachycudonia (Imai 1950). It has also been placed in the genus
Spathularia (Mains 1956).
Genetic studies (Ge
et al. 2014) suggest that
Pachycudonia can be considered a synonym of
Cudonia. No genetic analysis has been done on
Pachycudonia spathulata, but based on micro-features, there is no evidence to suggest it doesn't belong in
Cudonia.
Geographic range
This species is known from southern Oregon through the Sierra Nevada foothills, and eastern slopes of the coast range and around the northern portion of the Central Valley of California, south to Santa Cruz, County, California with two reports from northern British Columbia and two reports from northern Alaska. These northern reports may represent a different species as the habitat is very different from that noted in the more southern, and more abundant, reports; however they are retained in the assessment for now. It is probably under reported and more widespread in California and Oregon than currently known.
Population and Trends
The population remains poorly documented, however as this species has become better known, more subpopulations have been reported. The scarcity of reports probably has more to do with the fact that this species has not appeared in popular field guides.
Based on the habitat it is now known to occur in, this species is probably a lot more common than reported, and stable.
Population Trend: Stable
Habitat and Ecology
It is scattered to gregarious in duff under manzanita (
Arctostaphylos spp.) and Pacific Madrone (
Arbutus menziesii) in the Coast Ranges and Sierra Nevada Foothills. Its nutritional mode not known. Fruiting is in winter and early spring.
Threats
No specific threats have been identified with regards to this species.
Conservation Actions
No specific conservation actions have been identified with regards to this species at this time. Surveys for this species in suitable habitat are needed to clarify its distribution, population status and trends.
Use and Trade
No use/trade is known.
Source and Citation
Mueller, G.M. & Siegel, N. 2025. Pachycudonia spathulata. The IUCN Red List of Threatened Species 2025: e.T198481380A198487264. Accessed on 22 November 2025.